From 27c00b5accd58ea5dd280f2c8a63faa2469262be Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:16:11 -0500 Subject: [PATCH 1/9] Add more docstrings to types and its interface --- docs/src/features.md | 129 +++++++++++++++++++++++++++++ docs/src/naivefinder.md | 2 +- docs/src/simplecodingrule.md | 28 ------- src/types.jl | 152 ++++++++++++++++++++++++++--------- 4 files changed, 245 insertions(+), 66 deletions(-) create mode 100644 docs/src/features.md diff --git a/docs/src/features.md b/docs/src/features.md new file mode 100644 index 0000000..e2f2a97 --- /dev/null +++ b/docs/src/features.md @@ -0,0 +1,129 @@ +## The ORF features + +The `ORF` type is designed to be flexible and can store various types of information about the ORF. This versatility allows it to hold data such as the score of the ORF based on a scoring function, the sequence of the ORF, or even the translated amino acid sequence. For example, in the `NaiveFinder` method, the `score` subfield is utilized to store the score of the ORF obtained from the scoring function. This capability is possible because the `ORF` type not only captures structural details of the ORF, such as the range, strand, and frame, but also provides a convenient field called `Features` for additional information. + +```julia +phi = dna"GTGTGAGGTTATAACGCCGAAGCGGTAAAAATTTTAATTTTTGCCGCTGAGGGGTTGACCAAGCGAAGCGCGGTAGGTTTTCTGCTTAGGAGTTTAATCATGTTTCAGACTTTTATTTCTCGCCATAATTCAAACTTTTTTTCTGATAAGCTGGTTCTCACTTCTGTTACTCCAGCTTCTTCGGCACCTGTTTTACAGACACCTAAAGCTACATCGTCAACGTTATATTTTGATAGTTTGACGGTTAATGCTGGTAATGGTGGTTTTCTTCATTGCATTCAGATGGATACATCTGTCAACGCCGCTAATCAGGTTGTTTCTGTTGGTGCTGATATTGCTTTTGATGCCGACCCTAAATTTTTTGCCTGTTTGGTTCGCTTTGAGTCTTCTTCGGTTCCGACTACCCTCCCGACTGCCTATGATGTTTATCCTTTGAATGGTCGCCATGATGGTGGTTATTATACCGTCAAGGACTGTGTGACTATTGACGTCCTTCCCCGTACGCCGGGCAATAACGTTTATGTTGGTTTCATGGTTTGGTCTAACTTTACCGCTACTAAATGCCGCGGATTGGTTTCGCTGAATCAGGTTATTAAAGAGATTATTTGTCTCCAGCCACTTAAGTGAGGTGATTTATGTTTGGTGCTATTGCTGGCGGTATTGCTTCTGCTCTTGCTGGTGGCGCCATGTCTAAATTGTTTGGAGGCGGTCAAAAAGCCGCCTCCGGTGGCATTCAAGGTGATGTGCTTGCTACCGATAACAATACTGTAGGCATGGGTGATGCTGGTATTAAATCTGCCATTCAAGGCTCTAATGTTCCTAACCCTGATGAGGCCGCCCCTAGTTTTGTTTCTGGTGCTATGGCTAAAGCTGGTAAAGGACTTCTTGAAGGTACGTTGCAGGCTGGCACTTCTGCCGTTTCTGATAAGTTGCTTGATTTGGTTGGACTTGGTGGCAAGTCTGCCGCTGATAAAGGAAAGGATACTCGTGATTATCTTGCTGCTGCATTTCCTGAGCTTAATGCTTGGGAGCGTGCTGGTGCTGATGCTTCCTCTGCTGGTATGGTTGACGCCGGATTTGAGAATCAAAAAGAGCTTACTAAAATGCAACTGGACAATCAGAAAGAGATTGCCGAGATGCAAAATGAGACTCAAAAAGAGATTGCTGGCATTCAGTCGGCGACTTCACGCCAGAATACGAAAGACCAGGTATATGCACAAAATGAGATGCTTGCTTATCAACAGAAGGAGTCTACTGCTCGCGTTGCGTCTATTATGGAAAACACCAATCTTTCCAAGCAACAGCAGGTTTCCGAGATTATGCGCCAAATGCTTACTCAAGCTCAAACGGCTGGTCAGTATTTTACCAATGACCAAATCAAAGAAATGACTCGCAAGGTTAGTGCTGAGGTTGACTTAGTTCATCAGCAAACGCAGAATCAGCGGTATGGCTCTTCTCATATTGGCGCTACTGCAAAGGATATTTCTAATGTCGTCACTGATGCTGCTTCTGGTGTGGTTGATATTTTTCATGGTATTGATAAAGCTGTTGCCGATACTTGGAACAATTTCTGGAAAGACGGTAAAGCTGATGGTATTGGCTCTAATTTGTCTAGGAAATAACCGTCAGGATTGACACCCTCCCAATTGTATGTTTTCATGCCTCCAAATCTTGGAGGCTTTTTTATGGTTCGTTCTTATTACCCTTCTGAATGTCACGCTGATTATTTTGACTTTGAGCGTATCGAGGCTCTTAAACCTGCTATTGAGGCTTGTGGCATTTCTACTCTTTCTCAATCCCCAATGCTTGGCTTCCATAAGCAGATGGATAACCGCATCAAGCTCTTGGAAGAGATTCTGTCTTTTCGTATGCAGGGCGTTGAGTTCGATAATGGTGATATGTATGTTGACGGCCATAAGGCTGCTTCTGACGTTCGTGATGAGTTTGTATCTGTTACTGAGAAGTTAATGGATGAATTGGCACAATGCTACAATGTGCTCCCCCAACTTGATATTAATAACACTATAGACCACCGCCCCGAAGGGGACGAAAAATGGTTTTTAGAGAACGAGAAGACGGTTACGCAGTTTTGCCGCAAGCTGGCTGCTGAACGCCCTCTTAAGGATATTCGCGATGAGTATAATTACCCCAAAAAGAAAGGTATTAAGGATGAGTGTTCAAGATTGCTGGAGGCCTCCACTATGAAATCGCGTAGAGGCTTTGCTATTCAGCGTTTGATGAATGCAATGCGACAGGCTCATGCTGATGGTTGGTTTATCGTTTTTGACACTCTCACGTTGGCTGACGACCGATTAGAGGCGTTTTATGATAATCCCAATGCTTTGCGTGACTATTTTCGTGATATTGGTCGTATGGTTCTTGCTGCCGAGGGTCGCAAGGCTAATGATTCACACGCCGACTGCTATCAGTATTTTTGTGTGCCTGAGTATGGTACAGCTAATGGCCGTCTTCATTTCCATGCGGTGCACTTTATGCGGACACTTCCTACAGGTAGCGTTGACCCTAATTTTGGTCGTCGGGTACGCAATCGCCGCCAGTTAAATAGCTTGCAAAATACGTGGCCTTATGGTTACAGTATGCCCATCGCAGTTCGCTACACGCAGGACGCTTTTTCACGTTCTGGTTGGTTGTGGCCTGTTGATGCTAAAGGTGAGCCGCTTAAAGCTACCAGTTATATGGCTGTTGGTTTCTATGTGGCTAAATACGTTAACAAAAAGTCAGATATGGACCTTGCTGCTAAAGGTCTAGGAGCTAAAGAATGGAACAACTCACTAAAAACCAAGCTGTCGCTACTTCCCAAGAAGCTGTTCAGAATCAGAATGAGCCGCAACTTCGGGATGAAAATGCTCACAATGACAAATCTGTCCACGGAGTGCTTAATCCAACTTACCAAGCTGGGTTACGACGCGACGCCGTTCAACCAGATATTGAAGCAGAACGCAAAAAGAGAGATGAGATTGAGGCTGGGAAAAGTTACTGTAGCCGACGTTTTGGCGGCGCAACCTGTGACGACAAATCTGCTCAAATTTATGCGCGCTTCGATAAAAATGATTGGCGTATCCAACCTGCAGAGTTTTATCGCTTCCATGACGCAGAAGTTAACACTTTCGGATATTTCTGATGAGTCGAAAAATTATCTTGATAAAGCAGGAATTACTACTGCTTGTTTACGAATTAAATCGAAGTGGACTGCTGGCGGAAAATGAGAAAATTCGACCTATCCTTGCGCAGCTCGAGAAGCTCTTACTTTGCGACCTTTCGCCATCAACTAACGATTCTGTCAAAAACTGACGCGTTGGATGAGGAGAAGTGGCTTAATATGCTTGGCACGTTCGTCAAGGACTGGTTTAGATATGAGTCACATTTTGTTCATGGTAGAGATTCTCTTGTTGACATTTTAAAAGAGCGTGGATTACTATCTGAGTCCGATGCTGTTCAACCACTAATAGGTAAGAAATCATGAGTCAAGTTACTGAACAATCCGTACGTTTCCAGACCGCTTTGGCCTCTATTAAGCTCATTCAGGCTTCTGCCGTTTTGGATTTAACCGAAGATGATTTCGATTTTCTGACGAGTAACAAAGTTTGGATTGCTACTGACCGCTCTCGTGCTCGTCGCTGCGTTGAGGCTTGCGTTTATGGTACGCTGGACTTTGTGGGATACCCTCGCTTTCCTGCTCCTGTTGAGTTTATTGCTGCCGTCATTGCTTATTATGTTCATCCCGTCAACATTCAAACGGCCTGTCTCATCATGGAAGGCGCTGAATTTACGGAAAACATTATTAATGGCGTCGAGCGTCCGGTTAAAGCCGCTGAATTGTTCGCGTTTACCTTGCGTGTACGCGCAGGAAACACTGACGTTCTTACTGACGCAGAAGAAAACGTGCGTCAAAAATTACGTGCGGAAGGAGTGATGTAATGTCTAAAGGTAAAAAACGTTCTGGCGCTCGCCCTGGTCGTCCGCAGCCGTTGCGAGGTACTAAAGGCAAGCGTAAAGGCGCTCGTCTTTGGTATGTAGGTGGTCAACAATTTTAATTGCAGGGGCTTCGGCCCCTTACTTGAGGATAAATTATGTCTAATATTCAAACTGGCGCCGAGCGTATGCCGCATGACCTTTCCCATCTTGGCTTCCTTGCTGGTCAGATTGGTCGTCTTATTACCATTTCAACTACTCCGGTTATCGCTGGCGACTCCTTCGAGATGGACGCCGTTGGCGCTCTCCGTCTTTCTCCATTGCGTCGTGGCCTTGCTATTGACTCTACTGTAGACATTTTTACTTTTTATGTCCCTCATCGTCACGTTTATGGTGAACAGTGGATTAAGTTCATGAAGGATGGTGTTAATGCCACTCCTCTCCCGACTGTTAACACTACTGGTTATATTGACCATGCCGCTTTTCTTGGCACGATTAACCCTGATACCAATAAAATCCCTAAGCATTTGTTTCAGGGTTATTTGAATATCTATAACAACTATTTTAAAGCGCCGTGGATGCCTGACCGTACCGAGGCTAACCCTAATGAGCTTAATCAAGATGATGCTCGTTATGGTTTCCGTTGCTGCCATCTCAAAAACATTTGGACTGCTCCGCTTCCTCCTGAGACTGAGCTTTCTCGCCAAATGACGACTTCTACCACATCTATTGACATTATGGGTCTGCAAGCTGCTTATGCTAATTTGCATACTGACCAAGAACGTGATTACTTCATGCAGCGTTACCATGATGTTATTTCTTCATTTGGAGGTAAAACCTCTTATGACGCTGACAACCGTCCTTTACTTGTCATGCGCTCTAATCTCTGGGCATCTGGCTATGATGTTGATGGAACTGACCAAACGTCGTTAGGCCAGTTTTCTGGTCGTGTTCAACAGACCTATAAACATTCTGTGCCGCGTTTCTTTGTTCCTGAGCATGGCACTATGTTTACTCTTGCGCTTGTTCGTTTTCCGCCTACTGCGACTAAAGAGATTCAGTACCTTAACGCTAAAGGTGCTTTGACTTATACCGATATTGCTGGCGACCCTGTTTTGTATGGCAACTTGCCGCCGCGTGAAATTTCTATGAAGGATGTTTTCCGTTCTGGTGATTCGTCTAAGAAGTTTAAGATTGCTGAGGGTCAGTGGTATCGTTATGCGCCTTCGTATGTTTCTCCTGCTTATCACCTTCTTGAAGGCTTCCCATTCATTCAGGAACCGCCTTCTGGTGATTTGCAAGAACGCGTACTTATTCGCCACCATGATTATGACCAGTGTTTCCAGTCCGTTCAGTTGTTGCAGTGGAATAGTCAGGTTAAATTTAATGTGACCGTTTATCGCAATCTGCCGACCACTCGCGATTCAATCATGACTTCGTGATAAAAGATTGA" + +phiorfs = findorfs(phi, finder=NaiveFinder, minlen=75, scheme=lors) + +124-element Vector{ORF{4, NaiveFinder}}: + ORF{NaiveFinder}(9:101, '-', 3) + ORF{NaiveFinder}(100:627, '+', 1) + ORF{NaiveFinder}(223:447, '-', 1) + ORF{NaiveFinder}(248:436, '+', 2) + ORF{NaiveFinder}(257:436, '+', 2) + ORF{NaiveFinder}(283:627, '+', 1) + ORF{NaiveFinder}(344:436, '+', 2) + ORF{NaiveFinder}(532:627, '+', 1) + ORF{NaiveFinder}(636:1622, '+', 3) + ORF{NaiveFinder}(687:1622, '+', 3) + ORF{NaiveFinder}(774:1622, '+', 3) + ORF{NaiveFinder}(781:1389, '+', 1) + ORF{NaiveFinder}(814:1389, '+', 1) + ORF{NaiveFinder}(829:1389, '+', 1) + ORF{NaiveFinder}(861:1622, '+', 3) + ⋮ + ORF{NaiveFinder}(4671:5375, '+', 3) + ORF{NaiveFinder}(4690:4866, '+', 1) + ORF{NaiveFinder}(4728:5375, '+', 3) + ORF{NaiveFinder}(4741:4866, '+', 1) + ORF{NaiveFinder}(4744:4866, '+', 1) + ORF{NaiveFinder}(4777:4866, '+', 1) + ORF{NaiveFinder}(4806:5375, '+', 3) + ORF{NaiveFinder}(4863:5258, '-', 3) + ORF{NaiveFinder}(4933:5019, '+', 1) + ORF{NaiveFinder}(4941:5375, '+', 3) + ORF{NaiveFinder}(5082:5375, '+', 3) + ORF{NaiveFinder}(5089:5325, '+', 1) + ORF{NaiveFinder}(5122:5202, '-', 1) + ORF{NaiveFinder}(5152:5325, '+', 1) + ORF{NaiveFinder}(5164:5325, '+', 1) +``` + +In the example above we calculated a score using the `lors` scoring scheme (see [lors](https://github.com/camilogarciabotero/BioMarkovChains.jl/blob/533e53d97cf5951f1ca050454bce1423ec8d7c36/src/transitions.jl#L179) from the [BioMarkovChains.jl](https://camilogarciabotero.github.io/BioMarkovChains.jl/dev/) package). The score is stored in the `score` subfield of the `ORF` . + +All features can be accesed using a conviniente funciton called `features` that returns a `NamedTuple` with the features of the `ORF` and can be broadcasted to the entire collection of `ORF`s using the `.` syntax. + +```julia +features.(phiorfs) + +124-element Vector{@NamedTuple{score::Float64}}: + (score = -3.002461366087374,) + (score = -10.814621287968222,) + (score = -5.344187934894264,) + (score = -1.316724559874126,) + (score = -1.796631200562138,) + (score = -3.2651518608269856,) + (score = -1.4019264441082822,) + (score = -2.3192349590107475,) + (score = 5.055524446434241,) + (score = 2.7116397224896436,) + (score = 2.2564640592402165,) + (score = 1.777499581940097,) + (score = 2.3474811908011186,) + (score = 2.38568188352799,) + (score = 2.498608044469827,) + ⋮ + (score = -5.474837954151803,) + (score = 0.6909362932156138,) + (score = -5.900045211699447,) + (score = 1.2010656615619415,) + (score = 0.8541931309205604,) + (score = 2.7897961643147777,) + (score = -4.42890346770467,) + (score = -5.40624241726446,) + (score = -0.8080572222081075,) + (score = -5.571494087742448,) + (score = -4.882156920421228,) + (score = -5.639670353834974,) + (score = -0.8764121443326865,) + (score = -4.308687693802273,) + (score = -4.459423419810693,) +``` + +In this case the `lors` calculates the log odds ratio of the ORF sequence given two Markov models, one for the coding region and one for the non-coding region. The score is stored in the `score` field of the `NamedTuple` returned by the `features` function. By default the `lors` function return the base 2 logarithm of the odds ratio, so it is analogous to the bits of information that the ORF sequence is coding. + +The `NaiveFinder` method takes also `kwargs...` that are passed the the scoring function, so the `lors` function could be modified to calculate the inverse of the odds ratio by simply calling the models in the opposite order. + +```julia + invphiorfs = findorfs(phi, finder=NaiveFinder, minlen=75, scheme=lors, modela=ECOLINOCDS, modelb=ECOLICDS) + +features.(invphiorfs) + +124-element Vector{@NamedTuple{score::Float64}}: + (score = 3.0024613660873727,) + (score = 10.814621287968192,) + (score = 5.344187934894256,) + (score = 1.316724559874125,) + (score = 1.7966312005621363,) + (score = 3.265151860826984,) + (score = 1.401926444108279,) + (score = 2.319234959010748,) + (score = -5.055524446434286,) + (score = -2.711639722489663,) + (score = -2.256464059240233,) + (score = -1.7774995819401198,) + (score = -2.347481190801142,) + (score = -2.3856818835280125,) + (score = -2.498608044469844,) + ⋮ + (score = 5.474837954151802,) + (score = -0.6909362932156132,) + (score = 5.900045211699445,) + (score = -1.2010656615619424,) + (score = -0.8541931309205605,) + (score = -2.789796164314777,) + (score = 4.428903467704666,) + (score = 5.406242417264438,) + (score = 0.8080572222081076,) + (score = 5.571494087742444,) + (score = 4.882156920421224,) + (score = 5.639670353834972,) + (score = 0.8764121443326848,) + (score = 4.308687693802271,) + (score = 4.459423419810688,) +``` + +This is the same as the previous example but with the inverse of the odds ratio, meaning that the score is the the chance of the ORF sequence being non-coding given the two models. + diff --git a/docs/src/naivefinder.md b/docs/src/naivefinder.md index 075516b..62124c5 100644 --- a/docs/src/naivefinder.md +++ b/docs/src/naivefinder.md @@ -63,7 +63,7 @@ orfs = findorfs(seq, finder=NaiveFinder) Two other methods where implemented into `sequence` to get the ORFs in DNA or aminoacid sequences, respectively. They use the `findorfs` function to first get the ORFs and then get the correspondance array of `BioSequence` objects. ```julia -sequece.(orfs) +sequence.(orfs) 12-element Vector{NucSeq{4, DNAAlphabet{4}}} ATGCAACCCTGA diff --git a/docs/src/simplecodingrule.md b/docs/src/simplecodingrule.md index 5873845..1ccc84d 100644 --- a/docs/src/simplecodingrule.md +++ b/docs/src/simplecodingrule.md @@ -72,34 +72,6 @@ orfsdna = findorfs(seq, minlen=75, alternative_start=true) .|> sequence ``` -Now, we can score the sequences using the *log-odds ratio* score in the same line: - -```julia -orfsfeat = findorfs(seq, minlen=75, alternative_start=true, scheme=lors) .|> features - -20-element Vector{@NamedTuple{score::Float64}}: - (score = -2.5146325834372343,) - (score = -4.857592765476053,) - (score = -1.9986133020444345,) - (score = -3.4106894574555824,) - (score = -1.763485388728319,) - (score = 0.6825864481251348,) - (score = 0.21287161698917936,) - (score = -0.28187825646085224,) - (score = -1.373474082107631,) - (score = -4.273794970087796,) - (score = -2.3961559066784597,) - (score = -2.3663038090046142,) - (score = -0.8406863072332524,) - (score = 1.8013554455006733,) - (score = -2.0768031699080756,) - (score = -1.734088708668584,) - (score = -2.9820908143871194,) - (score = -3.072550585883162,) - (score = -2.712493281013948,) - (score = -2.0453354284951786,) -``` - Now the question is which of those sequences can we consider as coding sequences. We can use the `iscoding` predicate to check whether a sequence is coding or not based on the *log-odds ratio* decision rule: ```julia diff --git a/src/types.jl b/src/types.jl index d6a0998..2421607 100644 --- a/src/types.jl +++ b/src/types.jl @@ -6,11 +6,6 @@ export features, sequence, source export groupname, finder, frame, scheme, score, strand, STRAND_BOTH, STRAND_NEG, STRAND_POS, STRAND_NA #### Ribosome Binding Site (RBS) struct #### - -# seq[orf.first-bin01.offset.start:orf.first-1] - -# motifs = [dna"GGA", dna"GAG", dna"AGG"] - struct Features{K} fts::K end @@ -26,6 +21,12 @@ struct RBS end end +# seq[orf.first-bin01.offset.start:orf.first-1] + +# motifs = [dna"GGA", dna"GAG", dna"AGG"] + +####### End of RBS struct ####### + """ struct ORF{N,F} <: GenomicFeatures.AbstractGenomicInterval{F} @@ -42,6 +43,31 @@ The `ORF` struct represents an Open Reading Frame (ORF) in genomics. - `scheme::Union{Nothing,Function}`: The scheme used for the ORF. # Constructor + +```julia +ORF{N,F}( + groupname::String, + first::Int64, + last::Int64, + strand::Strand, + frame::Int, + features::Features, + scheme::Union{Nothing,Function} +) + +# Example + +A full instance `ORF` + +```julia +ORF{4,NaiveFinder}("seq01", 1, 33, STRAND_POS, 1, Features((score = 0.0,)), nothing) +``` + +A partial instance `ORF` + +```julia +ORF{NaiveFinder}(1:33, '+', 1) +``` """ struct ORF{N,F} <: GenomicFeatures.AbstractGenomicInterval{F} groupname::String @@ -86,7 +112,7 @@ function ORF{F}( strand::Char, frame::Int ) where {F<:GeneFinderMethod} - groupname = "unamedseq" + groupname = "unnamedsource" first = range.start last = range.stop features = Features(NamedTuple()) @@ -95,29 +121,6 @@ function ORF{F}( return ORF{4,F}(groupname, first, last, strand, frame, features, scheme) end -function groupname(i::ORF{N,F}) where {N,F} - return i.groupname -end - -function first(i::ORF{N,F}) where {N,F} - return i.first -end - -function last(i::ORF{N,F}) where {N,F} - return i.last -end - -function frame(i::ORF{N,F}) where {N,F} - return i.frame -end - -finder(i::ORF{N,F}) where {N,F} = F - -function scheme(i::ORF{N,F}) where {N,F} - return i.scheme -end - - """ sequence(i::ORF{N,F}) @@ -136,11 +139,90 @@ function sequence(i::ORF{N,F}) where {N,F} return i.strand == STRAND_POS ? @view(seq[i.first:i.last]) : reverse_complement(@view(seq[i.first:i.last])) #seq[i.first:i.last] end + +""" + source(i::ORF{N,F}) + +Get the source sequence associated with the given `ORF` object. + +# Arguments +- `i::ORF{N,F}`: The `ORF` object for which to retrieve the source sequence. + +# Returns +The source sequence associated with the `ORF` object. + +# Examples +```julia +seq = dna"ATGATGCATGCATGCATGCTAGTAACTAGCTAGCTAGCTAGTAA" +orfs = findorfs(seq) +source(orfs[1]) + +44nt DNA Sequence: +ATGATGCATGCATGCATGCTAGTAACTAGCTAGCTAGCTAGTAA +``` + +!!! warning + The `source` method works if the sequence is defined in the global scope. Otherwise it will throw an error. + For instance a common failure is to define a simple `ORF` that by defualt will have an "unnamedsource" as `groupname` + and then try to get the source sequence. + + ```julia + orf = ORF{NaiveFinder}(1:33, '+', 1) + source(orf) + + ERROR: UndefVarError: `unnamedsource` not defined + Stacktrace: + [1] source(i::ORF{4, NaiveFinder}) + @ GeneFinder ~/.julia/dev/GeneFinder/src/types.jl:192 + [2] top-level scope + @ REPL[12]:1 + ``` + +""" function source(i::ORF{N,F}) where {N,F} seqsymb = Symbol(i.groupname) return getfield(Main, seqsymb) end + +""" + features(i::ORF{N,F}) + +Extracts the features from an `ORF` object. + +# Arguments +- `i::ORF{N,F}`: An `ORF` object. + +# Returns +The features of the `ORF` object. + +""" +function features(i::ORF{N,F}) where {N,F} + return i.features.fts +end + +function groupname(i::ORF{N,F}) where {N,F} + return i.groupname +end + +function first(i::ORF{N,F}) where {N,F} + return i.first +end + +function last(i::ORF{N,F}) where {N,F} + return i.last +end + +function frame(i::ORF{N,F}) where {N,F} + return i.frame +end + +finder(i::ORF{N,F}) where {N,F} = F + +function scheme(i::ORF{N,F}) where {N,F} + return i.scheme +end + function score(i::ORF{N,F}) where {N,F} return i.features.fts[:score] end @@ -149,14 +231,14 @@ function length(i::ORF{N,F}) where {N,F} return length(sequence(i)) end -function features(i::ORF{N,F}) where {N,F} - return i.features.fts -end - function strand(i::ORF{N,F}) where {N,F} return i.strand end +function metadata(i::ORF{N,F}) where {N,F} + return features(i) +end + function Base.show(io::IO, i::ORF{N,F}) where {N,F} if get(io, :compact, false) print(io, "ORF{$(finder(i))}($(leftposition(i)):$(rightposition(i)), '$(strand(i))', $(frame(i)))") #{$(typeof(finder(i)))} $(score(i)) @@ -165,10 +247,6 @@ function Base.show(io::IO, i::ORF{N,F}) where {N,F} end end -function metadata(i::ORF{N,F}) where {N,F} - return features(i) -end - ## Ideas for Gene struct # struct CDS <: AbstractGene From adb624253814decb0b47478f92105cbda351da72 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:17:46 -0500 Subject: [PATCH 2/9] Add a new section analysing scoring ORFs to docs --- docs/src/features.md | 9 ++++++++- 1 file changed, 8 insertions(+), 1 deletion(-) diff --git a/docs/src/features.md b/docs/src/features.md index e2f2a97..186aae9 100644 --- a/docs/src/features.md +++ b/docs/src/features.md @@ -125,5 +125,12 @@ features.(invphiorfs) (score = 4.459423419810688,) ``` -This is the same as the previous example but with the inverse of the odds ratio, meaning that the score is the the chance of the ORF sequence being non-coding given the two models. +This is the same as the previous example but with the inverse of the odds ratio, meaning that the score is the the chance of the ORF sequence being non-coding given the same two models used before (i.e. ECOLICDS and ECOLINOCDS). Now we can even analyse how is the distribution of the scores in the ORFs. + +```julia +using UnicodePlots +``` + + + From 4baaef2b4fa8a46909c43b487067ec7f3d769265 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:28:04 -0500 Subject: [PATCH 3/9] Update documentation with new section on scoring ORFs --- docs/src/api.md | 7 ------- 1 file changed, 7 deletions(-) diff --git a/docs/src/api.md b/docs/src/api.md index c712e4e..9e2fc71 100644 --- a/docs/src/api.md +++ b/docs/src/api.md @@ -30,13 +30,6 @@ Modules = [GeneFinder] Pages = ["algorithms/naivefinder.jl"] ``` - - ## Writing ORFs to files ```@autodocs From 1e07d03801bddf3bb70153c447b6a08c233e0331 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:29:16 -0500 Subject: [PATCH 4/9] refactor: rename _locationiterator function in naivefinder.jl --- src/algorithms/naivefinder.jl | 2 +- 1 file changed, 1 insertion(+), 1 deletion(-) diff --git a/src/algorithms/naivefinder.jl b/src/algorithms/naivefinder.jl index f09f4af..05a501c 100644 --- a/src/algorithms/naivefinder.jl +++ b/src/algorithms/naivefinder.jl @@ -5,7 +5,7 @@ abstract type GeneFinderMethod end struct NaiveFinder <: GeneFinderMethod end """ - locationiterator(sequence::NucleicSeqOrView{DNAAlphabet{N}}; alternative_start::Bool=false) where {N} + _locationiterator(sequence::NucleicSeqOrView{DNAAlphabet{N}}; alternative_start::Bool=false) where {N} This is an iterator function that uses regular expressions to search the entire ORF (instead of start and stop codons) in a `LongSequence{DNAAlphabet{4}}` sequence. It uses an anonymous function that will find the first regularly expressed ORF. Then using this anonymous function it creates an iterator that will apply it until there is no other CDS. From 314bbfdaefb33c9f8fc7752c567bd4a842cf739e Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:30:30 -0500 Subject: [PATCH 5/9] docs: Add new section on scoring ORFs to documentation --- docs/make.jl | 3 ++- 1 file changed, 2 insertions(+), 1 deletion(-) diff --git a/docs/make.jl b/docs/make.jl index b166f41..7f7d98b 100644 --- a/docs/make.jl +++ b/docs/make.jl @@ -18,6 +18,7 @@ makedocs(; pages = [ "Home" => "index.md", "Finding ORFs" => "naivefinder.md", + "Scoring ORFs" => "features.md", "A Simple Coding Rule" => "simplecodingrule.md", "Wrtiting ORFs In Files" => "iodocs.md", "API" => "api.md", @@ -26,4 +27,4 @@ makedocs(; warnonly = true, ) -deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "main") \ No newline at end of file +deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "scoredocs") \ No newline at end of file From 738de61000751d55bd4383bf4a4312d8990f6358 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:32:00 -0500 Subject: [PATCH 6/9] docs: remove blank space --- docs/src/simplecodingrule.md | 1 - 1 file changed, 1 deletion(-) diff --git a/docs/src/simplecodingrule.md b/docs/src/simplecodingrule.md index 1ccc84d..d4d1d34 100644 --- a/docs/src/simplecodingrule.md +++ b/docs/src/simplecodingrule.md @@ -129,7 +129,6 @@ orfs[iscoding.(orfsdna)] Or in a single line using another genome sequence: ```julia - phi = dna"GTGTGAGGTTATAACGCCGAAGCGGTAAAAATTTTAATTTTTGCCGCTGAGGGGTTGACCAAGCGAAGCGCGGTAGGTTTTCTGCTTAGGAGTTTAATCATGTTTCAGACTTTTATTTCTCGCCATAATTCAAACTTTTTTTCTGATAAGCTGGTTCTCACTTCTGTTACTCCAGCTTCTTCGGCACCTGTTTTACAGACACCTAAAGCTACATCGTCAACGTTATATTTTGATAGTTTGACGGTTAATGCTGGTAATGGTGGTTTTCTTCATTGCATTCAGATGGATACATCTGTCAACGCCGCTAATCAGGTTGTTTCTGTTGGTGCTGATATTGCTTTTGATGCCGACCCTAAATTTTTTGCCTGTTTGGTTCGCTTTGAGTCTTCTTCGGTTCCGACTACCCTCCCGACTGCCTATGATGTTTATCCTTTGAATGGTCGCCATGATGGTGGTTATTATACCGTCAAGGACTGTGTGACTATTGACGTCCTTCCCCGTACGCCGGGCAATAACGTTTATGTTGGTTTCATGGTTTGGTCTAACTTTACCGCTACTAAATGCCGCGGATTGGTTTCGCTGAATCAGGTTATTAAAGAGATTATTTGTCTCCAGCCACTTAAGTGAGGTGATTTATGTTTGGTGCTATTGCTGGCGGTATTGCTTCTGCTCTTGCTGGTGGCGCCATGTCTAAATTGTTTGGAGGCGGTCAAAAAGCCGCCTCCGGTGGCATTCAAGGTGATGTGCTTGCTACCGATAACAATACTGTAGGCATGGGTGATGCTGGTATTAAATCTGCCATTCAAGGCTCTAATGTTCCTAACCCTGATGAGGCCGCCCCTAGTTTTGTTTCTGGTGCTATGGCTAAAGCTGGTAAAGGACTTCTTGAAGGTACGTTGCAGGCTGGCACTTCTGCCGTTTCTGATAAGTTGCTTGATTTGGTTGGACTTGGTGGCAAGTCTGCCGCTGATAAAGGAAAGGATACTCGTGATTATCTTGCTGCTGCATTTCCTGAGCTTAATGCTTGGGAGCGTGCTGGTGCTGATGCTTCCTCTGCTGGTATGGTTGACGCCGGATTTGAGAATCAAAAAGAGCTTACTAAAATGCAACTGGACAATCAGAAAGAGATTGCCGAGATGCAAAATGAGACTCAAAAAGAGATTGCTGGCATTCAGTCGGCGACTTCACGCCAGAATACGAAAGACCAGGTATATGCACAAAATGAGATGCTTGCTTATCAACAGAAGGAGTCTACTGCTCGCGTTGCGTCTATTATGGAAAACACCAATCTTTCCAAGCAACAGCAGGTTTCCGAGATTATGCGCCAAATGCTTACTCAAGCTCAAACGGCTGGTCAGTATTTTACCAATGACCAAATCAAAGAAATGACTCGCAAGGTTAGTGCTGAGGTTGACTTAGTTCATCAGCAAACGCAGAATCAGCGGTATGGCTCTTCTCATATTGGCGCTACTGCAAAGGATATTTCTAATGTCGTCACTGATGCTGCTTCTGGTGTGGTTGATATTTTTCATGGTATTGATAAAGCTGTTGCCGATACTTGGAACAATTTCTGGAAAGACGGTAAAGCTGATGGTATTGGCTCTAATTTGTCTAGGAAATAACCGTCAGGATTGACACCCTCCCAATTGTATGTTTTCATGCCTCCAAATCTTGGAGGCTTTTTTATGGTTCGTTCTTATTACCCTTCTGAATGTCACGCTGATTATTTTGACTTTGAGCGTATCGAGGCTCTTAAACCTGCTATTGAGGCTTGTGGCATTTCTACTCTTTCTCAATCCCCAATGCTTGGCTTCCATAAGCAGATGGATAACCGCATCAAGCTCTTGGAAGAGATTCTGTCTTTTCGTATGCAGGGCGTTGAGTTCGATAATGGTGATATGTATGTTGACGGCCATAAGGCTGCTTCTGACGTTCGTGATGAGTTTGTATCTGTTACTGAGAAGTTAATGGATGAATTGGCACAATGCTACAATGTGCTCCCCCAACTTGATATTAATAACACTATAGACCACCGCCCCGAAGGGGACGAAAAATGGTTTTTAGAGAACGAGAAGACGGTTACGCAGTTTTGCCGCAAGCTGGCTGCTGAACGCCCTCTTAAGGATATTCGCGATGAGTATAATTACCCCAAAAAGAAAGGTATTAAGGATGAGTGTTCAAGATTGCTGGAGGCCTCCACTATGAAATCGCGTAGAGGCTTTGCTATTCAGCGTTTGATGAATGCAATGCGACAGGCTCATGCTGATGGTTGGTTTATCGTTTTTGACACTCTCACGTTGGCTGACGACCGATTAGAGGCGTTTTATGATAATCCCAATGCTTTGCGTGACTATTTTCGTGATATTGGTCGTATGGTTCTTGCTGCCGAGGGTCGCAAGGCTAATGATTCACACGCCGACTGCTATCAGTATTTTTGTGTGCCTGAGTATGGTACAGCTAATGGCCGTCTTCATTTCCATGCGGTGCACTTTATGCGGACACTTCCTACAGGTAGCGTTGACCCTAATTTTGGTCGTCGGGTACGCAATCGCCGCCAGTTAAATAGCTTGCAAAATACGTGGCCTTATGGTTACAGTATGCCCATCGCAGTTCGCTACACGCAGGACGCTTTTTCACGTTCTGGTTGGTTGTGGCCTGTTGATGCTAAAGGTGAGCCGCTTAAAGCTACCAGTTATATGGCTGTTGGTTTCTATGTGGCTAAATACGTTAACAAAAAGTCAGATATGGACCTTGCTGCTAAAGGTCTAGGAGCTAAAGAATGGAACAACTCACTAAAAACCAAGCTGTCGCTACTTCCCAAGAAGCTGTTCAGAATCAGAATGAGCCGCAACTTCGGGATGAAAATGCTCACAATGACAAATCTGTCCACGGAGTGCTTAATCCAACTTACCAAGCTGGGTTACGACGCGACGCCGTTCAACCAGATATTGAAGCAGAACGCAAAAAGAGAGATGAGATTGAGGCTGGGAAAAGTTACTGTAGCCGACGTTTTGGCGGCGCAACCTGTGACGACAAATCTGCTCAAATTTATGCGCGCTTCGATAAAAATGATTGGCGTATCCAACCTGCAGAGTTTTATCGCTTCCATGACGCAGAAGTTAACACTTTCGGATATTTCTGATGAGTCGAAAAATTATCTTGATAAAGCAGGAATTACTACTGCTTGTTTACGAATTAAATCGAAGTGGACTGCTGGCGGAAAATGAGAAAATTCGACCTATCCTTGCGCAGCTCGAGAAGCTCTTACTTTGCGACCTTTCGCCATCAACTAACGATTCTGTCAAAAACTGACGCGTTGGATGAGGAGAAGTGGCTTAATATGCTTGGCACGTTCGTCAAGGACTGGTTTAGATATGAGTCACATTTTGTTCATGGTAGAGATTCTCTTGTTGACATTTTAAAAGAGCGTGGATTACTATCTGAGTCCGATGCTGTTCAACCACTAATAGGTAAGAAATCATGAGTCAAGTTACTGAACAATCCGTACGTTTCCAGACCGCTTTGGCCTCTATTAAGCTCATTCAGGCTTCTGCCGTTTTGGATTTAACCGAAGATGATTTCGATTTTCTGACGAGTAACAAAGTTTGGATTGCTACTGACCGCTCTCGTGCTCGTCGCTGCGTTGAGGCTTGCGTTTATGGTACGCTGGACTTTGTGGGATACCCTCGCTTTCCTGCTCCTGTTGAGTTTATTGCTGCCGTCATTGCTTATTATGTTCATCCCGTCAACATTCAAACGGCCTGTCTCATCATGGAAGGCGCTGAATTTACGGAAAACATTATTAATGGCGTCGAGCGTCCGGTTAAAGCCGCTGAATTGTTCGCGTTTACCTTGCGTGTACGCGCAGGAAACACTGACGTTCTTACTGACGCAGAAGAAAACGTGCGTCAAAAATTACGTGCGGAAGGAGTGATGTAATGTCTAAAGGTAAAAAACGTTCTGGCGCTCGCCCTGGTCGTCCGCAGCCGTTGCGAGGTACTAAAGGCAAGCGTAAAGGCGCTCGTCTTTGGTATGTAGGTGGTCAACAATTTTAATTGCAGGGGCTTCGGCCCCTTACTTGAGGATAAATTATGTCTAATATTCAAACTGGCGCCGAGCGTATGCCGCATGACCTTTCCCATCTTGGCTTCCTTGCTGGTCAGATTGGTCGTCTTATTACCATTTCAACTACTCCGGTTATCGCTGGCGACTCCTTCGAGATGGACGCCGTTGGCGCTCTCCGTCTTTCTCCATTGCGTCGTGGCCTTGCTATTGACTCTACTGTAGACATTTTTACTTTTTATGTCCCTCATCGTCACGTTTATGGTGAACAGTGGATTAAGTTCATGAAGGATGGTGTTAATGCCACTCCTCTCCCGACTGTTAACACTACTGGTTATATTGACCATGCCGCTTTTCTTGGCACGATTAACCCTGATACCAATAAAATCCCTAAGCATTTGTTTCAGGGTTATTTGAATATCTATAACAACTATTTTAAAGCGCCGTGGATGCCTGACCGTACCGAGGCTAACCCTAATGAGCTTAATCAAGATGATGCTCGTTATGGTTTCCGTTGCTGCCATCTCAAAAACATTTGGACTGCTCCGCTTCCTCCTGAGACTGAGCTTTCTCGCCAAATGACGACTTCTACCACATCTATTGACATTATGGGTCTGCAAGCTGCTTATGCTAATTTGCATACTGACCAAGAACGTGATTACTTCATGCAGCGTTACCATGATGTTATTTCTTCATTTGGAGGTAAAACCTCTTATGACGCTGACAACCGTCCTTTACTTGTCATGCGCTCTAATCTCTGGGCATCTGGCTATGATGTTGATGGAACTGACCAAACGTCGTTAGGCCAGTTTTCTGGTCGTGTTCAACAGACCTATAAACATTCTGTGCCGCGTTTCTTTGTTCCTGAGCATGGCACTATGTTTACTCTTGCGCTTGTTCGTTTTCCGCCTACTGCGACTAAAGAGATTCAGTACCTTAACGCTAAAGGTGCTTTGACTTATACCGATATTGCTGGCGACCCTGTTTTGTATGGCAACTTGCCGCCGCGTGAAATTTCTATGAAGGATGTTTTCCGTTCTGGTGATTCGTCTAAGAAGTTTAAGATTGCTGAGGGTCAGTGGTATCGTTATGCGCCTTCGTATGTTTCTCCTGCTTATCACCTTCTTGAAGGCTTCCCATTCATTCAGGAACCGCCTTCTGGTGATTTGCAAGAACGCGTACTTATTCGCCACCATGATTATGACCAGTGTTTCCAGTCCGTTCAGTTGTTGCAGTGGAATAGTCAGGTTAAATTTAATGTGACCGTTTATCGCAATCTGCCGACCACTCGCGATTCAATCATGACTTCGTGATAAAAGATTGA" filter(x -> iscoding(sequence(x), η=1e-10) && length(x) > 100, findorfs(phi)) From f772e7e2b6fce50b40a582d76fec331f34d98325 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:32:56 -0500 Subject: [PATCH 7/9] chore: Update deploydocs function with mainbranch parameter --- docs/make.jl | 2 +- 1 file changed, 1 insertion(+), 1 deletion(-) diff --git a/docs/make.jl b/docs/make.jl index 7f7d98b..3f82259 100644 --- a/docs/make.jl +++ b/docs/make.jl @@ -27,4 +27,4 @@ makedocs(; warnonly = true, ) -deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "scoredocs") \ No newline at end of file +deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "scoredocs", mainbranch = "main") \ No newline at end of file From e228b93f4c6775d2f2b119a8c284023ada764023 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 15:42:09 -0500 Subject: [PATCH 8/9] chore: Update deploydocs function with mainbranch parameter --- docs/make.jl | 2 +- 1 file changed, 1 insertion(+), 1 deletion(-) diff --git a/docs/make.jl b/docs/make.jl index 3f82259..3bf4fdd 100644 --- a/docs/make.jl +++ b/docs/make.jl @@ -27,4 +27,4 @@ makedocs(; warnonly = true, ) -deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "scoredocs", mainbranch = "main") \ No newline at end of file +deploydocs(; repo = "https://github.com/camilogarciabotero/GeneFinder.jl", devbranch = "main") \ No newline at end of file From 8b59fe03411f63f6cf843edc38817cc681e42a37 Mon Sep 17 00:00:00 2001 From: =?UTF-8?q?Camilo=20Garc=C3=ADa?= Date: Wed, 3 Jul 2024 17:42:50 -0500 Subject: [PATCH 9/9] feat: Add analysis of ORF scoring to documentation --- docs/src/assets/lors-lamda.png | Bin 0 -> 211321 bytes docs/src/features.md | 94 +++++++++++++++++---------------- 2 files changed, 48 insertions(+), 46 deletions(-) create mode 100644 docs/src/assets/lors-lamda.png diff --git 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-4.459423419810693,) ``` -In this case the `lors` calculates the log odds ratio of the ORF sequence given two Markov models, one for the coding region and one for the non-coding region. The score is stored in the `score` field of the `NamedTuple` returned by the `features` function. By default the `lors` function return the base 2 logarithm of the odds ratio, so it is analogous to the bits of information that the ORF sequence is coding. +## Analysing Lamda ORFs -The `NaiveFinder` method takes also `kwargs...` that are passed the the scoring function, so the `lors` function could be modified to calculate the inverse of the odds ratio by simply calling the models in the opposite order. +In this case the `lors` calculates the log odds ratio of the ORF sequence given two Markov models (by default: ECOLICDS and ECOLINOCDS), one for the coding region and one for the non-coding region. The score is stored in the `score` field of the `NamedTuple` returned by the `features` function. By default the `lors` function return the base 2 logarithm of the odds ratio, so it is analogous to the bits of information that the ORF sequence is coding. -```julia - invphiorfs = findorfs(phi, finder=NaiveFinder, minlen=75, scheme=lors, modela=ECOLINOCDS, modelb=ECOLICDS) - -features.(invphiorfs) - -124-element Vector{@NamedTuple{score::Float64}}: - (score = 3.0024613660873727,) - (score = 10.814621287968192,) - (score = 5.344187934894256,) - (score = 1.316724559874125,) - (score = 1.7966312005621363,) - (score = 3.265151860826984,) - (score = 1.401926444108279,) - (score = 2.319234959010748,) - (score = -5.055524446434286,) - (score = -2.711639722489663,) - (score = -2.256464059240233,) - (score = -1.7774995819401198,) - (score = -2.347481190801142,) - (score = -2.3856818835280125,) - (score = -2.498608044469844,) - ⋮ - (score = 5.474837954151802,) - (score = -0.6909362932156132,) - (score = 5.900045211699445,) - (score = -1.2010656615619424,) - (score = -0.8541931309205605,) - (score = -2.789796164314777,) - (score = 4.428903467704666,) - (score = 5.406242417264438,) - (score = 0.8080572222081076,) - (score = 5.571494087742444,) - (score = 4.882156920421224,) - (score = 5.639670353834972,) - (score = 0.8764121443326848,) - (score = 4.308687693802271,) - (score = 4.459423419810688,) -``` - -This is the same as the previous example but with the inverse of the odds ratio, meaning that the score is the the chance of the ORF sequence being non-coding given the same two models used before (i.e. ECOLICDS and ECOLINOCDS). Now we can even analyse how is the distribution of the scores in the ORFs. +Now we can even analyse how is the distribution of the ORFs' scores as a function of their lengths compared to random sequences. ```julia -using UnicodePlots -``` - - +lambda = fasta_to_dna("test/data/NC_001416.1.fasta")[1] + +lambaorfs = findorfs(lambda, finder=NaiveFinder, minlen=100, scheme=lors) + +lamdascores = score.(lambaorfs) +lambdalengths = length.(lambaorfs) + +## get some random sequences of variable lengths +vseqs = LongDNA[] +for i in 1:708 + push!(vseqs, randdnaseq(rand(100:1000))) +end + +## get the lengths and scores of the random generated sequences +randlengths = length.(vseqs) +randscores = lors.(vseqs) + +## plot the scores as a function of the lengths +using CairoMakie + +f = Figure() +ax = Axis(f[1, 1], xlabel="Length", ylabel="Log-odds ratio (Bits)") + +scatter!(ax, + randlengths, + randscores, + marker = :circle, + markersize = 6, + color = :black, + label = "Random sequences" +) +scatter!(ax, + lambdalengths, + lambdascores, + marker = :rect, + markersize = 6, + color = :blue, + label = "Lambda ORFs" +) + +axislegend(ax) + +f +``` +![](assets/lors-lamda.png) \ No newline at end of file